Finn Russell, says Scotland need to “up it” if they are to progress in the Rugby World Cup, despite recording a 45-17 win over Tonga in Nice on Sunday.

He said that while the number of tries scored by seven different players is great, he believes that four, five, or six opportunities were missed.

In training and games, we need to be harder on ourselves and be a lot better when we get to the finishing zone five metres out. While it’s great that the boys are scoring, we need to do a lot better there. Our performance next week will be frustrating and we need to improve if we want to advance. We’re happy we got the bonus point, but we need to improve.”

Aside from South Africa, Ireland beat the defending champions in Pool B this weekend.

“It would have been ideal for us if South Africa had won, but we know what needs to be done now,” said Russell.

“I don’t think it changes much for us. After the South Africa game, we knew we had to beat Ireland in Paris. It’s good to know what we had to do.”

Just 10 days after receiving a call up to join the Scotland World Cup squad, Stuart McInally has been forced to withdraw with a neck injury.

Dave Cherry, another Edinburgh hooker who slipped on hotel stairs and suffered a concussion, requested the former captain’s help.

Johnny Matthews will now fill the third hooker position for Glasgow Warriors.

When players are rested for next weekend’s meeting with Romania in Lille, Matthews, 30, may well provide support for George Turner and Ewan Ashman.